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Fig. 5. The distribution of integumentary types in coelurosaurian theropod dinosaurs and inferred distribution of plumage patterning. Protofeather-like appendages appeared at the base of Coelurosauria, if not earlier. Color patterns reported in the tail of the compsognathid Sinosauropteryx indicate that among-feather color patterns may have also appeared at this stage. Pinnate feathers and within-feather color patterns first appear in Maniraptora, observed in the striped pinnate tail feathers of the oviraptorosaur Caudipteryx and the troodontid Anchiornis huxleyi.

DOI: 10.1126/science.1186290

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Uploaded on February 5, 2010